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  • The article has not been previously published, and has not been sent to another journal for consideration.
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  • The person making the submission has been authorized by all the article authors to submit and act as their spokesperson in front of the journal during the review, editing and publishing processes.
  • I have consulted and applied the journal's Research data policy.
  • The text adheres to the length, format, references, citation of figures, tables and equations (if applicable), and bibliography requirements outlined in the journal guidelines.
  • Each of the authors has been identified including the following data:
    - Given name (in full form) and family name(s).
    - Email contact address.
    - Country of professional activity.
    - Institutional affiliation.
    - Open Researcher and Contributor Identifier (ORCID).

Author Guidelines

Download HERE the Good Practice Code in PDF
Download HERE the Authorship Form in PDF


Anales del Jardín Botánico de Madrid (AJBM) is a journal published by the Consejo Superior de Investigaciones Científicas (CSIC) in electronic format and with free access on the web (

AJBM publishes original scientific research articles and short notes in the field of Botany (taxonomy, systematics, anatomy, morphology, evolution, phylogenetics, genomics, ecology, ecophysiology, reproductive biology, biogeography, conservation, and ethnobotany), and in relation to organisms from cyanobacteria to vascular plants.

Authors of research involving the participation of human subjects, should consider the incorporation of sex or gender in the experimental design, analysis of data, and interpretation of results.

Sending drafts for publication

Drafts, either in English or Spanish, should be sent through the journal's webpage. The drafts should consist of:
― A single MS Word file (.doc, .docx or .rtf) including the text and legends of tables and figures at its end. When possible, authors should use the provided templates for Articles or Short Notes.
― Tables, each one in a single MS Word file (.doc, .docx or .rtf).
― Figures, each one in a single file (see below information about file formats for figures). All them should be submitted in a single compressed .zip or .rar file.
― The text should follow the Recommendations for the use of inclusive and non-sexist language.
― The authorship form signed by each author should be sent along with the draft.

General format

Use Times New Roman font with size 12. Do not justify right margin. Number text lines and pages. Line numbers should be removed in the final accepted version of the manuscript. All the illustrations (photographs, drawings, maps, and graphics) are treated as figures, are mentioned in the text and are numbered consecutively according to the order in which they are cited.

Organization of the draft

Contents of the first page

Title: up to a maximum of 120 characters. When appropriate, it must include one or two suprageneric ranks for the taxon or taxa treated. For example, “Synopsis of Chionanthus (Oleaceae) in Colombia and bordering countries and description of a new species”. Do not include the authors of scientific names.

List of authors: Include their corresponding postal addresses, email address of corresponding author/s, and ORCID identifiers ( ORCID identifiers are mandatory for all authors; manuscript missing ORCIDs will not be processed. Each author will be associated to a superscript number that also precedes each of the aforementioned attributes. The author for correspondence is identified with a superscript asterisk.

Abbreviated title: up to a maximum of 50 characters (not required for short notes).

Abstract: up to a maximum of 200 words and its translation. The Abstract should not contain bibliographic references. Commonly accepted abbreviations may be used, such as h, min, s, mL, L, mg/L, °C, d, wk, mo, ITS, RPD, RFLP, rDNA, 18S etc.

Keywords: a maximum of five keywords in alphabetical order.

Resumen: Abstract in Spanish.

Palabras clave: keywords in Spanish.

Contents of the subsequent pages

The main body of research articles includes the following sections: “INTRODUCTION”, “MATERIAL AND METHODS”, “RESULTS”, “DISCUSSION”, “ACKNOWLEDGMENTS” (optional), and “REFERENCES”. After the last section, include legends of tables and figures in order. For taxonomic treatments and checklists, the results and the discussion may be combined as “RESULTS AND DISCUSSION”.

Title of sections are in regular capital letters. Subsections are in lowercase italics with initial capital letter.

In the case of short notes, the main body does not follow the structure of the body of a research article (above). A few subtitles can be used if necessary; see examples of short notes published in recent issues of AJBM.

Bibliographic references

In the text, they should be cited as follows: “Pau (1903)”, “Pau (1903: 273)”, “(Pau 1903)”, “(Pau 1903: 273)” or “Pau (1903, 1907)”. Multiple references should be included in parentheses in chronological order: “(Pau 1903; Font Quer 1934; Maas & Craigie 1964; Cornejo & al. 2011)”. References published in the same year will be distinguished by lowercase letters: “(Sánchez 2010a, 2010b; Fernández & Cogollo 2016a, 2016b)”. When the citations refer to several articles, with more than two authors, whose first author is the same they should be distinguished as follows: “(Sánchez & al. 2001a, 2001b)”, “(Sánchez & al. 2001b)”. In the list of references use the same lowercase letters as in the text.

In the section "REFERENCES", required for both research articles and short notes, the bibliographical references should be listed alphabetically. For works with a single author, use the first surname of the author (if it is the same, the author’s first name is used). Those references sharing the same first author should be arranged in chronological order. If the work is signed by two authors, and the first is the same, they will be sorted alphabetically by the last name of the second author (or by the initial letter of the name if they coincide); if they were the same, they will be ordered chronologically and the same lowercase letter code (a, b, c, ...) will be used for their citation in the text. When the work is signed by three or more authors the order will be chronological. All authors will be included; the expression "& al." is not used in this section. The titles of the publications will not be abbreviated in this section (unlike in the citation of protologues) and should be those recognized in the works Botanico-Periodicum-Huntianum (Lawrence & al. 1968; Bridson 1991, 2004,, as well as from Taxonomic Literature (Stafleu & Cowan 1976-1988, and its supplements, when they include them. Only published works should be included. Unpublished data or articles, or observations made by colleagues, will not be included in the format of a reference. The DOI (Digital Object Identifier, should be provided for each reference when possible.

See the examples prepared for each type of publication at the end of this document.

Figures and legends

Use TIF, TIFF, JPG, or JPEG file formats for halftones and bitmaps. For vector-based images use SVG, EPS, AI, or PDF. The minimum final resolution for images in color or grayscale (halftones, bitmaps) should be 350 dpi; for botanical line drawings in black and white use 1200 dpi (file size can be reduced using LZW compression). Resolution issues do not apply to vector-based images.

All figures must contain scales. Identify each part of composite figures (plates) with a lowercase letter. In graphs, the coordinate axes should be labelled.

Legends should be self-explanatory. Indicate the name of the taxon, collector and collection number, herbarium code, the value of the scales, etc. when necessary. Letters indicating different photos or parts of a figure should be in small caps in bold (a, b, c, d,). See example below for additional formatting and punctuation.

Legend examples:

Fig. 22. Geranium californicum G.N.Jones & F.L.Jones: a, habit; b, sheet; c, inflorescence; d, flower; e, staminal filament; f, fruit; g, mericarpo [a, d-f, Hall and Chandler 224 (UC); b, c, g, Ertter 6665. (MA 234567); scales: a-c, f = 1 cm; d, e = 1 mm; g = 0.5 mm].

Fig. 9. Quararibea villanuevae Fern.Alonso sp. nov.: a, details of the fruit and the cupular calyx that accompanies the fruit; b, detail of the open pyrene and the seed; c-d, detail of the seed showing the testa with the funicular scar and the mass of the cotyledon [Villanueva & Tamayo 2446 (TOLI); photos: B. Villanueva; scales: a, b, c, d = 1 cm].


The nomenclature of scientific names should agree with the provisions of the current International Nomenclature Code.

The authors of scientific names must be included at the first mention of the name in the body of the text (also in figures and tables; and in the abstract if nomenclatural issues are discussed). Authorities are not included in the title of the article, nor in keywords. The genus name in a binomial should not be abbreviated at the first mention in a paragraph or as the first word in a sentence; they can be abbreviated elsewhere in the body of the text. Authorities should be cited using the abbreviations proposed by Brummit & Powell (1992) and their updates in the IPNI database, see

In taxonomic treatments, the bibliographic references to the protologue or the name of the publication of each scientific name should be abbreviated according to Botanico-Periodicum-Huntianum (Hunt 2004, for journal titles and Taxonomic Literature ed. 2 (Stafleu & Cowan 1976-1988, and its supplements for book titles. The publication should be included in the REFERENCES section.


Centaurea cadevallii Pau in Butll. Inst. Catalana Hist. Nat. 6: 46 (1906).
Centaurea polycephala sensu Willk. in Willk. & Lange, Prodr. Fl. Hispan. 2: 156 (1865), non Jord.


Cadevall J. 1906. Excursionisme botanich. Butlletí de la Institució Catalana d'Història Natural 6: 41–46.
Willkomm H.M. & Lange J.M.C. 1865. Prodromus florae Hispanicae 2: 139. Sumtibus E. Schweizerbart (E. Koch), Stuttgart.

The specimens used for taxonomic revisions, as well as for chromosome counting or any analysis, or for the extraction of DNA, must be deposited in herbaria accessible to the public, especially the type material. Herbaria should be indicated in the text as acronyms, following Index Herbariorum (; Thiers 2019). The DNA sequences must be accompanied by their registration number in the GenBank database (

In taxonomic treatments, the name of each accepted taxon should be written in bold italic font and numbered. (Indicate subspecific names by adding a lowercase letter in alphabetical order). The synonyms should be grouped by homotypic blocks in independent paragraphs.

As for the typographical conventions adopted for the reproduction of herbarium labels and handwritten notes, a simple bar ( / ) will be used to symbolize line breaks; a double bar ( // ) to separate different labels present in the same sheet; quotes ( “ ” ) to reproduce what can not be interpreted or formatted in a satisfactory way or that, deliberately, the author wants to reproduce in a literal way (verbatim); brackets ( [ ] ) to distinguish information that has been interpreted or inferred from the label text, such as localities and translations of the text; and ellipses ( ... ) to represent a part of an illegible word, word or passage in a handwritten text. Likewise, when such literal representation of a label is intended, the italics will be used for the transcription of the handwritten parts, the bold italics for the transcription of words or underlined passages and the round or regular type is used for the transcription of printed parts.

See the examples at the end of this document, where you will also find a table with the most commonly used abbreviations.

Identification keys should be dichotomous, with numbered entries, without indentation and should include only accepted taxa of the treatment (if other taxa of lower rank are also accepted, these may or may not be integrated in the general key). See an example of a dichotomous identification key at the end of this document.

For material cited in the text, use the following format, with countries arranged alphabetically: COUNTRY. First order administrative division: Locality, coordinates, elevation (m), date of collection, collector and collector number (herbarium codes). See an example at the end of the document.

An index of collections may be added as an appendix at the end of the manuscript, after legends and tables. Entries should be ordered alphabetically by collector's last name and by collection number. The format is as follows: Collector and collector number, number of the accepted taxon with which it corresponds in the treatment. See an example of index of collections at the end of this document.


Relevant checklists that provide a sufficient number of corological novelties will be admitted for publication. The presentation of the studied or cited material will be adjusted to the examples shown at the end of this document. Bibliographic citations will include all the collection data present in the original publication, expanded or clarified if possible.


The sentences or headings of the general text should not begin with an abbreviation (apart from taxonomic blocks).

The numbers from one to nine should be written out (one, two, three, etc.) unless they correspond to a measure, a range or an indicator, or the number of organs in the morphological descriptions.

When mentioning figures in the text, use the following style: Fig. 1; Figs. 1, 2; Figs. 13; Figs. 2, 56, 89; Figs. 1c, 2b; Fig. 2ce; Fig. 1a, b.

Units should be abbreviated using standards proposed by the International System of Units.

See the list of common abbreviations at the end of this document.

References mentioned above

Bridson G.D.R. (ed.) 1991. B-P-H / S: Botanical-Periodicum-Huntianum / Supplementum. Hunt Institute for botanical documentation, Pittsburg.

Bridson G.D.R. (ed.) 2004. BPH-2: Periodicals with Botanical Content. Vols. 12. Hunt Institute for botanical documentation, Pittsburg.

Lawrence G.H.M., Günther Buchheim A.F., Daniels G.S. & Dolezal H. (eds.) 1968. B-P-H: Botanico-Periodicum-Huntianum. Hunt Institute for botanical documentation, Pittsburg.

Stafleu R.A. & Cowan R.S. 1976-1988. Taxonomic Literature: A Selective Guide to Botanical Publications and Collections with Dates, Commentaries and Types ed. 2. Vols. 94, 98, 105, 110, 112, 115116. Utrech, Antwerp, The Hague and Boston.

Stearn W.T. 1992. Botanical Latin: History, Grammar, Syntax, Terminology, and Vocabulary ed. 4. Timber Press, Portland.

Thiers B. 2019 [continuously updated]. Index Herbariorum: A global directory of public herbal and associated staff. New York Botanical Garden's Virtual Herbarium, New York.

Turland N.J., Wiersema J.H., Barrie F.R., Greuter W., Hawksworth D.L., Herendeen P.S., Knapp S., Kusber W.-H., Li D.-Z., Marhold K., May T.W., McNeill J., Monro A.M., Prado J., Price M.J. & Smith G.F. (eds.) 2018. International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. [Regnum Vegetabile 159.] Koeltz Botanical Books, Glashütten.


― Article published in a journal:

Cuatrecasas J. 1958. Aspectos de la vegetación natural de Colombia. Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales 10: 221–268.

Aedo C., Medina L., Barberá P. & Fernández-Albert M. 2015. Extinctions of vascular plants in Spain. Nordic Journal of Botany 33: 83–100.

Helm M., Schmid M., Hierl G., Terneus K., Tan L., Lottspeich F., Kieliszewski M.J. & Gietl C. 2008. KDEL-tailed cysteine endopeptidases involved in programmed cell death, intercalation of new cells and dismantling of extensin scaffolds. American Journal of Botany 95: 1049–1062.

― Article published in a special volume (journals):

Santos Guerra A. 1996. Notas corológicas III: Adiciones florísticas y nuevas localidades para la flora canaria. In Aedo A. & Muñoz Garmendia F. (eds.), Volume honoring Manuel Laínz. Anales del Jardín Botánico de Madrid 54: 445‒448.

― Book:

Peterson R.L., Massicotte H.B. & Melville L.H. 2004. Mycorrhizas: Anatomy and Cell Biology. CABI Publishing Series, New York.

Esau K. 1976. Plant Anatomy ed. 2. Wiley, New York.

Kuntze O. 1943. Revisio Generum Plantarum 1: 35–37. Arthur Felix, Leipzig.

Jonker F.P. 1891. Triuridaceae. In Pulle A. (ed.), Flora of Suriname (Netherlands Guyana) vol. 1. J.H. de Bussy, Amsterdam.

― Book that belongs to a series:

Walter H. 1977. Zonas de vegetación y clima. Manuales flexibles de Ciencias Biológicas. Ediciones Omega, Barcelona.

― Chapter or article in a book:

Smith C.A., Alberts J.J. & Bond J. 2001. Pyrus L. In Castroviejo S. (ed.), Flora iberica 2: 313–406. Editorial CSIC, Madrid.

Gentry A.H. 1995. Patterns of Diversity and Floristic Composition in Neotropial Montane Forest. In Churchill S.P., Balslev H., Forero E. & Luteyn J.L. (eds.), Biodiversity and Conservation of Neotropical Montane Forests: 103‒126. The New York Botanical Garden, New York.

― Book with several volumes or parts:

Rico E., Crespo M.B., Quintanar A., Herrero A. & Aedo C. (eds.). 2013. Liliaceae-Agavaceae. In Castroviejo S. (coord.), Flora iberica vol. 20. Editorial CSIC, Madrid.

― Article in congress book:

D'Arcy W.G. 1991. The Solanaceae since 1976, with a review of its biogeography. In Hawkes G.J., Lester R.N., Nee M. & Estrada N. (eds.), Solanaceae III: Taxonomy, Chemistry, Evolution: 75–137. Proceedings of Third International Conference on Solanaceae. Kew Publishing, Kew.

― Ph.D. Thesis or technical report:

Černá K. 2010. Spatial variability and ecology of phytobenthic algal assemblages in peat bogs. Ph.D.dissertation, Charles University of Prague, Prague.

Conkle M.T., Hodgskiss P.D. & Hunter S.C. 1982. Starch gel electrophoresis of conifer seeds: A laboratory manual. USDA Forest Service General Technical Report PSW-64, Pacific Southwest Forest and Range Experiment Station, Berkeley.

― Websites:

Watson L. & Dallwitz M.J. 1992. The families of flowering plants: descriptions, illustrations, identification, and information retrieval. Website: [accessed: 2 Apr. 2017].

― Presentations in scientific meetings and posters:

Vila-Viçosa C., Gonçalves J., Honrado J., Garcia C., Almeida R., Vázquez F.M. & Lomba A. 2018, March 20–24. Bridging Temperate and Mediterranean Regions: insights from marcescent forests in Iberian Peninsula. Climate Change Biogeography. Article presented at the meeting of the International Biogeographical Society, Evora, Portugal.

Adams-Labonte S.K. 2012, August. Daytime impairment due to college students’ technology use during sleep: Similarities to sleep apnea. Poster session presented at the meeting of the American Psychological Association, Orlando, Florida.

― Symposia:

Krinsky-McHale S.J., Zigman W.B. & Silverman W. 2012, August. Are neuropsychiatric symptoms markers of prodromal Alzheimer’s disease in adults with Down syndrome? In Zigman W.B. (Chair), Predictors of mild cognitive impairment, dementia, and mortality in adults with Down syndrome. Symposium conducted at the meeting of the American Psychological Association, Orlando, Florida.

― Meetings or conventions in blogs:

Mills K.I. 2012. Why do people hurt themselves? [Blog post]. Website: [accessed: 25 Jul. 2012].


― Name accepted:

8. Teucrium rivasii Rigual ex Greuter & Burdet, Willdenowia 15: 81 (1985); T. rivasii Rigual, Fl. Alicante: 344 (1972), nom. inval.; T. rivasii var. glabrescens Rigual, Fl. Alicante: 344 (1972), nom. inval.; T. buxifolium subsp. rivasii (Greuter & Burdet) M.B.Crespo, Mateo & Güemes, Bol. Soc. Brot. ser. 2, 63: 125 (1990); T. buxifolium var. rivasii (Greuter & Burdet) M.B.Crespo, Mem. Soc. Brot. 29: 121 (1993). Original citation: “In fissuris prope Novelda.” [sec. Rigual (1972: 334)]. Type: [Spain] Monte Agudo [Monteagudo de Novelda], 3 May 1954, A. Rigual s.n. (holotype: ABH 23804 photo!, ABH 23807 photo!).

― Name accepted with heterotypic synonym:

3. Centaurea hanrii Jord., Observ. Pl. Nouv. 5: 70, pl. 4B (1847); C. polycephala sensu Willk., Prodr. Fl. Hispan. 2: 156 (1865), non Jordan (1847); C. coerulescens subsp. hanrii (Jord.) Bonnier, Fl. Ill. France 6: 43 (1923). Original citation: “(...) par lui à la Sainte-Beaume près Toulon (Var).”. Type: [Francia] Ste. Baume (Var), 10 Jul. 1846, H. Hanry s.n. (lectotype designated by López & Devesa (2010: 123): LY-Bonaparte s.n.!).

C. cadevallii Pau, Butll. Inst. Catalana Hist. Nat. 6: 46 (1906). Original citation: “In nemorosis sterilibus agrorum marginibus Valldoreix, prope Sant Gugat del Vallés.” Type: [España] In nemorosis sterilibus agrorum marginibus Valldoreix, prope San Cugat del Vallés, J. Cadevall s.n. (lectotype designated by López & Devesa (2010: 122): MA 135357 [upper left specimen] photo!; isolectotypes: BC-Sennen s.n.n.v., MA 135358 photo!).

― Subspecies accepted:

6. Narcissus triandrus L., Sp. Pl. ed. 2: 416 (1762); Ganymedes triandrus (L.) Haw., Suppl. Pl. Succ.: 130 (1819); Queltia triandra (L.) M. Roem., Fam. Nat. Syn. Monogr. 4: 203 (1847). Original citation: “Habitat in Pyrenaeis.”. Type: Icon (lectotype designated by Barra & López (1982: 70): Clus., Exot. Libri: [20] (1605)).

6a. Narcissus triandrus subsp. triandrus

N. reflexus Brot., Fl. Lusit. 1: 550 (1804). Original citation: [Spain] “(...) in montosis Gerez, circa Amarante, et alibi in Duriminia.” Type: Not localized.

N. calathinus auct., non Linnaeus (1762).

6b. Narcissus triandrus subsp. pallidulus (Graells) Rivas Goday, Veg. Fl. Guadiana: 710 (1964); N. pallidulus Graells, Indic. Pl. Nov.: 9 (1854), bason. Original citation: “Crescit in ruprestibus regionis submontanae montium Carpetanorum: ad alt. 2500-4000'; etiam inveni in nemorosis Argandae aliisque Castellae.”. Type: [Spain, Madrid] Silla de Felipe II prima vere, M.P. Graells s.n. (lectotype designated by Aedo (2010: 138): MA 591812!; isolectotypes: BM n.v., GOET n.v., MACB 56584 foto!, MPU n.v., OXF n.v., P!).

― Only one subspecies accepted:

2. Bellevalia dubia subsp. hackelii (Freyn) Feinbrun, Palestine J. Bot., Jerusalem Ser. 1: 348 (1940) [“Hackeli”]; B. hackelii Freyn, Oesterr. Bot. Z. 27: 289 (1877), bason.; Hyacinthus dubius auct. Original citation: “Lusitaniae in collibus argilloso-lusitanico s. majo 1876 legit E. Hackel.”. Type: [Portugal] Not localized.

― Name on a label:

7. Stachys brachyclada De Noé in Balansa, Pl. Algérie no. 623 (1852), in sched. [“brachyoclada”]; S. hirta var. parviflora De Noé ex Bory & Durieu, Expl. Sci. Algérie, Bot. 1, pl. 65 (1848), nom. subst. Original citation: “Champs en friche avoisinant la rade de Mer-el-Kébir” [sec. F. de Noé in Balansa, Pl. Algérie no. 623 (1852), in sched.]. Type: [Argelia] Champs en friche avoisinant la rade de Mer-el-Kébir, 20 Feb. 1852, B. Balansa s.n. (syntypes: G 00424778 photo!, MPU 020651 photo!, MPU 020652 photo!, MPU 020653 photo!).


3. Petiole 6–18(20) mm; leaf blades with 5–10 pairs of axillary domatia on the veins of the underside; axillary inflorescences 5–10 cm; corolla 13–17 mm; mature fruit 30–40 × 25–30 mm, subglobose, black .................. 3. Ch. colonchensis
– Petiole (12)15–30(35) mm; leaf blades without axillary domatia on the veins of the underside; axillary inflorescences 8–16(20) cm; corolla less than 10 mm; ripe fruit 45–50 × 42–47 mm, ellipsoid, purple ...... 8. Ch. megistocarpus
4. Leaves (15)20–30 cm; leaf blades with 10–20 pairs of lateral veins; inflorescences of 10 cm or more in length ...................................................... 5
– Leaves usually up to 15 cm; leaf blades with up to 10 pairs of lateral veins; inflorescences up to 10 cm ........................................ 7
5. Leaves with petioles 2–5 mm; foliar blades with 18–20 pairs of lateral veins, without domatia on the underside ........................................ 12. Ch. wurdackii
– Leaves with petioles 13–35 mm; leaf blades with 12–18 pairs of lateral veins, with domatia on the underside ............................................................ 6


Additional specimens examined.—FRANCE. Bouches-du-Rhone: Marseille, 3 May 1914 (P), (casual alien). Corse-du-Sud: Bonifacio, May 1917 (P). ITALY. Basilicata: Potenza, 3 Jun. 1928 (FI) ; Potenza, Monte Caruso, 1841 (NAP). Calabria: Catanzaro, Tiriolo, 15 Jun. 1899 (FI); Reggio Calabria, Capo d’Armi, 3 May 1824 (NAP); Vibo Valentia, 25 May 2009 (W). Campania: Avellino, Villamaina, 20 May 1832 (NAP); tra Sacco e Teggiano [locus classicus of var. montana!], 1000 m, 23 Jun.2001 (Herb.EDG)*; Felitto, 250 m, 30 May 2002 (Herb.EDG).

Additional specimens examined.—VENEZUELA. Mérida: Dtto. Tovar, 37.4 km beyond beginning of dirt portion of road (at turn off in Guayabal) to San Isidro Alto and El Molino on SE facing slope of steep valley above Hacienda Betania, 08°18’N, 71°36’W, 2850 m, 02 Dec. 1983, A.L. Weitzman & C. Sobrevila 34 (MERF s.n., VEN 203690). Táchira: Dtto. Uribante, 13 km S of El Portachuelo, on small road to Pregonero off Bailadores-La Grita road; heavily disturbed cloud forest along road; 08°07’N, 71°53’W, 2650 m, 27 Nov. 1983, A.L. Weitzman & C. Sobrevila 29 (MERF s.n., MY 91023); idem, A.L. Weitzman & C. Sobrevila 31 (MERF s.n., VEN 203898).


Ekman E.L. 13630 (14); Escobar M. & Santa J. 252 (14), 355 (14), 403 (17), 404 (17); Espinosa R. 19 (29); Felippone F. 720 (25), 837 (25), 839 (25), 839 (25), 960 p.p. (25), 990 (25), 2439 (25); Flores M. 308 (29); Florschütz P.A. 3974 (14), 4166 (14); Frahm J.-P. 824021 (4), 824071 (17); Frey T.C. 3072 & Frey E.M. (14).

The number in parentheses corresponds to the one that the accepted taxon takes in the treatment.


We include below a list with abbreviations commonly used in Botany, as they are used in AJBM. If not found here, preferably follow Stearn (1992).

Abbildung (illustration) Abb.
alii (others) al.
apud (in the publication of) ap.
auctorum (of the authors) auct.
basonym basón.
circiter (approximately), circa (around) c.
confer (compare with, see) cf.
combinatio (combination) comb.
cultivated cult.
cultivarietas (cultivar, taxonomic context) cv.
diameter of the trunk or bole at breast height (1.3 m above ground level) DBH
descriptio (description) descr.
editio (edition), editor (Editor) ed.
exceptus (excepted) exc.
exclusus (excluded) excl.
form (taxonomic rank) f.
family (taxonomic rank) fam.
fascicle fasc.
figure fig.
floruit (flourished), flower (in the citation of specimens, able to be accompanied by the symbols ♂, ♀ and ⚥) fl.
fruit (in the citation of specimens) fr.
genus (taxonomic rank) g.
group (taxonomic context) gr.
hybridus (hybrid) hybrid.
iconography ic.
ibidem (in the same place) ibid.
id est (namely) i. e.
inclusus (included) incl.
indicatio locotypica (original citation of locality) ind. loc.
unpublished (articles, data, results, etc.) inéd.
in erratis (in the errata of the work) in err.
in schedula (in a herbarium label) in sched.
plate lám.
loco citato (cited place) loc. cit.
locus classicus (type locality) loc. class.
number no.
manuscript ms.
nomen alternativum (alternative name) nom. alt.
nomen ambiguum (ambiguous name) nom. ambig.
nomen conservandum (name whose conservation has been accepted) nom. cons.
nomen conservandum propositum (name whose conservation has been proposed) nom. cons. prop.
nomen dubius (dubious name) nom. dub.
nomen illegitimum (illegitimate name) nom. illegit.
nomen invalidum (invalid name) nom. inval.
nomen nudum (nude name) nom. nud.
nomen provisorium (tentative name) nom. prov.
nomen rejiciendum (rejected name) nom. rejic.
nomen rejiciendum propositum (name proposed to be rejected) nom. rejic. prop.
nomen substitutum (replaced name) nom. subst.
nomen superfluum (superfluous name) nom. superfl.
nothosubspecies (hybrid subspecies) nothosubsp.
novus (new) nov.
non vidi (I have not seen it) n. v.
order (taxonomic context) ord.
orthographia conservanda (spelling whose conservation has been accepted) orth. cons.
pro maxima parte (so the most part does) p. max. p.
page p.
pages pp.
personal comment pers. comm.
plate pl.
prope (near, next to) pr.
sensu lato (in a broad sense) s.l.
sine anno (collections) s. a.
sine numero (collections, drawings, etc.) s. n.
sensu stricto (strictly speaking) s. str.
secundum (according to) sec.
sectio (section, taxonomic rank) sect.
series (series, taxonomic rank) ser.
meters above the sea level m a.s.l.
species (taxonomic rank) sp.
subclass (taxonomic rank) subcl.
subform (taxonomic rank) subf.
subfamily (taxonomic rank) subfam.
subgenus (taxonomic rank) subg.
subsection (taxonomic rank) subsect.
subspecies (taxonomic rank) subsp.
subvariety (taxonomic rank) subvar.
superfluus (superfluous) superfl.
synonymon substitutum (replaced synonym) syn. subst.
tabula (plate, illustration) tab.
Tafel (plate, illustration) Taf.
typus exclusus (excluding the type) typ. excl.
verbi gratia (for example) v. gr.
variety (taxonomic rank) var.
volume vol.
vidi (I have seen it) !


Below you will find instructions for the following procedures:

- How to register at Anales del Jardín Botánico de Madrid

- How to send an original manuscript to Anales del Jardín Botánico de Madrid

- How to check a revision report and submit a revised version of a manuscript


How to register at Anales del Jardín Botánico de Madrid

1.- On the journal's website, click on the link "Register":


2.- On the registration page you will have to fill in all the fields (the "Family name" field is optional, although highly recommended) using lower case only for your email, username and password. After reading the Privacy Statement you will need to consent to the collection and storage of your data and complete the Captcha. Receiving notifications of new publications and notices is optional.


3.- You will receive a message with a link to activate your account at the provided email address. Once activated, you will be able to log in to your account with the credentials you created.


How to send an original manuscript to Anales del Jardín Botánico de Madrid

1.- On the magazine's website, log in by clicking on "Login":


2.- Enter your username and password. Once you return to the magazine's homepage, click on the "Make a Submission" button:

Before submitting, please review the "Submission Preparation Checklist" and read the "Author Guidelines", the "Copyright Notice" and the "Privacy Statement". When you have all the required documents ready, click on the link "Make a new submission".

In subsequent logins, you will be taken to your user dashboard. There press the ‘New Submission’ button to submit a new article. You will be shown any articles received and in process under the ‘My Queue’ tab):


3.- As you will see on the new submissions page, submitting a manuscript is a four-stage process, in addition to a final section with information on follow-up:

3.1.- "1. Start"

At this stage you must select the language in which the article is written and the section of the journal in which you think it would fit, you must indicate that you have prepared all the items on the "Submission requirements" list and, optionally, you may send comments to the editor. You must comply with the terms of the Copyright Statement and the collection and storage of your data as the author of the article in accordance with the Privacy Statement of the Consejo Superior de Investigaciones Científicas (CSIC).

Then press the "Save and continue" button.


3.2.- "2. Upload submission"

In this phase, the files that make up the article and its additional documentation will be uploaded to the management module.

There are three parts to upload each file. In the first one, we will select which component of the article we are going to contribute and we will upload the corresponding file. Once uploaded, click on the "Continue" button. In the second part we will see the file metadata, with the possibility to edit them, but we will click "Continue" without making this edition.

In the third part, select "Add Another File", going back to "1. Upload file" and, without modifying the default option "This is not a revision of an existing file", select the new "Article Component" identifying what it is and uploading the file. Click "Continue" until you reach, once again, step "3. Confirm". This process must be repeated until all the files have been sent, and only when all the material has been uploaded should the "Complete" button be selected.

If after "Completing" the submission we realise that we have forgotten to upload a file, we can do so by selecting the "Upload File" button located in the upper right-hand corner of the "2. Upload Submission" tab:

Once all the submission files have been uploaded, we will press the "Save and continue" button.


3.3.- "3. Enter Metadata"

In this phase the author will enter the metadata of the article according to the journal guidelines. These are:

- Title in Spanish and English. If the article is written in another language, it will be introduced first this language and then in English.

- Summary or abstract in the same languages as the previous item.

- List of contributors. Although it was not necessary when registering as a journal user, in this section it is compulsory for authors to have their ORCID identifier and affiliation correctly indicated. If necessary, the information of additional authors will be added using the link "Add contributor":

- Keywords. The article keywords will be inserted in both languages. The entire list cannot be copied, must be entered one term at a time pressing "Enter" after each one.

- Funding data. The entities that have supported the research published in the article must be indicated. After selecting "Add funder", the name of the funder should be inserted again, which will trigger an internal search that will return the institution standardised name and DOI. If the institution does not have a DOI, it will not be able to register in this field. After entering the grant numbers, click on "Save".

To finish this phase of entering metadata, click on the "Save and continue" button.


3.4.- "4. Confirmation" y "5. Next steps"

In this last phase we will confirm the submission metadata recording linked to the uploaded files. Before clicking on the "Finish Submission" button, we can go back to the previous phases and review the information and files provided to check that they are all correct.

Once we click on the "Finish Submission" button, the article will be sent to the journal and its staff will contact you to continue with the process, as indicated in the "5. Next Steps" section.


How to check a revision report and submit a revised version of a manuscript

Once your submission has been reviewed, the journal's staff will send you a review report. Once received, you must log in to the journal and, in the "Submissions" section of your dashboard, you will be able to check that your submission is in the Review phase and, if the editorial staff has requested, whether it is necessary to make any modifications or revisions to the manuscript:

By clicking on the title of your submission, you will be taken to the workflow of your submission and you will be able to check the information related to the its review. You will find the notifications that the editorial staff has sent you, the attachments that the reviewers may have attached and, if requested, the possibility to provide a new version of the manuscript with the requested modifications by clicking on the "Upload file" button:


Research data policy

We recommend authors depositing data obtained from the research developed for the preparation of their article in repositories of recognized prestige, specific to the discipline or of a generalist nature. In any case, it must be a FAIR repository (Findable, Accessible, Interoperable, Reusable), preferably in open access.

There are several repositories destined to conserve and disseminate concrete data such as results of surveys, observations, interviews, simulations, automatically collected data, samples, models ... If necessary, authors can consult the Registry of Research Data Repositories re3data taking into account that each repository has its own deposit rules.

Those CSIC authors who would like to deposit their datasets in Digital.CSIC may do so by following these guidelines. They can use the Servicio de Archivo Delegado made available by the Technical Office of DIGITAL.CSIC and the Red de Bibliotecas CSIC.

DIGITAL.CSIC generates DOIs for datasets and associated software and is certified as data repository in re3data and Repository Finder. More information at Política de datos en Digital.CSIC.

If the author has deposited datasets in a repository, he should mention it in the article providing a brief description of the type of data deposited, the name and URL of the repository, the identification code and the data of the license for use and distribution. This information must be included at the end of the article, immediately before the bibliographic listing, under the heading "Data availability".

Download HERE the Good Practice Code in PDF
Download HERE the Authorship Form in PDF

Privacy Statement

The Spanish National Research Council (CSIC) has a record of data processing activities. Data collected through this form will be incorporated and processed in the “Gestión de las actividades de producción y distribución de las publicaciones del CSIC” treatment activity of Editorial CSIC, in order to manage the requested service. It is the responsibility of Editorial CSIC to manage this record. If you wish to exercise your rights, please contact us through the contact address Vitruvio, 8, 28006 Madrid, Spain, e-mail address Data processing is legitimized by the consent of the affected. The data may not be transferred to third parties except in the cases provided for in current regulations on the protection of personal data. You have the right to file a claim with the Spanish Data Protection Agency. You have the right to withdraw your consent. In the event that you wish -or want to exercise the rights of access, deletion, rectification, limitation or portability- you can do so through the following form. You can also contact the CSIC Officer for Data Protection via email: